Why do horses shake their heads?

What triggers a horse to flip his head uncontrollably, sometimes to the point of endangering him and his rider?

The amazing thing about horses is how such large, powerful animals can be so sensitive and aware of the slightest sensation, such as a fly on its back or face. Rippling of skin or an occasional head shake is a normal response to the tickling trigger of nerve endings. But, there are times when a horse can’t stop shaking or tossing its head to a seemingly in-apparent sensation; such incessant behaviour is known as head shaking.

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